Introduction 3 "Orsten"-type fossils 3 History of phosphatocopine research 3 Prior assumptions on the systematics of Phosphatocopina. 3 Soft parts and new perspectives about the phytogeny4 Scope and aims 4 Material and methods 4 Material 4 General information 4 Statistical data 5 Stratigraphy 6 Methods 7 Isolation of specimens (not done for this work) 7 Preparation for SEM 8 Measurements 8 Terminology 8 Systematics 10 Crustacea Brunnich, 1772 13 Phylogenetic analysis 13 Results 13 Hesslandona Milller, 1964. 15 Hesslandona abdominalis Hinz-Schallreuter, 1998 16 Hesslandona reichi Hinz-Schallreuter, 1993 16 Hesslandona unisulcata Milller, 1982. 16 Hesslandona necopina Milller, 1964. 58 Hesslandona kinnekullensis Muller, 1964 67 Hesslandona trituberculata (Lochman 8t Hu, 1960) Rushton, 1978 71 Hesslandona ventrospinata Grundel in Grundel 8c Buchholz, 1981 77 Hesslandona suecica n. sp 85 Hesslandona angustata n. sp 89 Hesslandona curvispina n. sp 93 Hesslandona toreborgensis n.

sp 104 Trapezilites Hinz-Schallreuter, 1993 106 Trapezilites minimus (Kummerow, 1931) Hinz-Schallreuter, 1993 106 Waldoria Griindel in Griindel 8c Buchholz, 1981 113 Waldoria rotundata Griindel in Griindel 8c Buchholz, 1981. 115 Veldotron Grundel in Griindel 8c Buchholz, 1981 121 Veldotron bratteforsa (Muller, 1964) Hinz-Schallreuter, 1993 123 Falites Muller, 1964 128 Falites fala Muller, 1964 131 Vestrogothia Muller, 1964 137 Vestrogothia spinata Muller, 1964 139 Discussion 145 Comparative morphology'' 145 Phylogenetic analysis of Phosphatocopina 159 General remarks 159 Reconstructed phytogeny of Phosphatocopina 160 Phosphatocopina ("Phosphatocopida sp." + Euphosphatocopina new name 160 Phosphatocopida sp 165 Euphosphatocopina new taxon 165 Vestrogothia spinata 168 NN1 ( Falites fala + Hesslandonina) 169 Falites fala 169 Hesslandonina (Cyclotronidae + Hesslandonidae) 170 Cyclotronidae (Veldotron bratteforsa + Cyclotron lapworthi ) 171 Veldotron bratteforsa 171 Cyclotron lapworthi 172 Waldoria rotundata 172 Hesslandonidae ( Trapezilites minimus + Hesslandona ) 173 Trapezilites minimus 173 Hesslandona 174 Hesslandona unisulcata 174 Dorsospinata new taxon 174 NN3 ( Hesslandona kinnekullensis +NN4) 175 Hesslandona kinnekullensis 175 NN4 ( Hesslandona toreborgensis n. sp., H. angustata n. sp. + H.

suecica n. sp.) 177 Hesslandona toreborgensis n. sp 177 Hesslandona suecica n. sp 177 Hesslandona angustata n. sp 178 NN5 ( Hesslandona ventrospinata + NN6) 178 Hesslandona ventrospinata . 179 NN6 ( Hesslandona trituberculata + NN7) 179 Hesslandona trituberculata .179 NN7 ( Hesslandona necopina + H.

curvispina ) 179 Hesslandona necopina 180 Hesslandona curvispina n. sp 180 Consequences of the proposed phytogeny of Phosphatocopina 180 Crustacean phytogeny 182 Ground pattern of Crustacea 182 The Ground pattern of Labrophora Siveter, Waloszek 8e Williams (2003) 185 Ground pattern of Phosphatocopina 190 Ground pattern of Eucrustacea 191 Conclusions 194 Future investigations and prospects 194 Acknowledgements 194 References 195 Appendix A - Data for the phylogenetic analysis 203 PAUP settings 203 List of all coded characters 203 Character matrix 206 Appendix B -- Synonymy of Phosphatocopine taxa 209 Possible Phosphatocopine taxa 209 Possible Phosphatocopines, described in open nomenclature 237.